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Pray excuse this long rambling note, and believe me, my dear sir, yours very sincerely, C. DARWIN.

Will you be so good as to present my respectful compliments to Sir W.

Hooker.

[Referring to Sir J.D. Hooker's work on the Galapagos Flora, my father wrote in 1846:

"I cannot tell you how delighted and astonished I am at the results of your examination; how wonderfully they support my assertion on the differences in the animals of the different islands, about which I have always been fearful."Again he wrote (1849):--"I received a few weeks ago your Galapagos papers (These papers include the results of Sir J.D. Hooker's examination of my father's Galapagos plants, and were published by the Linnean Society in 1849.), and I have read them since being here. I really cannot express too strongly my admiration of the geographical discussion: to my judgment it is a perfect model of what such a paper should be; it took me four days to read and think over. How interesting the Flora of the Sandwich Islands appears to be, how I wish there were materials for you to treat its flora as you have done the Galapagos. In the Systematic paper I was rather disappointed in not finding general remarks on affinities, structures, etc., such as you often give in conversation, and such as De Candolle and St. Hilaire introduced in almost all their papers, and which make them interesting even to a non-Botanist."

"Very soon afterwards [continues Sir J.D. Hooker] in a letter dated January 1844, the subject of the 'Origin of Species' was brought forward by him, and I believe that I was the first to whom he communicated his then new ideas on the subject, and which being of interest as a contribution to the history of Evolution, I here copy from his letter":--]

CHARLES DARWIN TO J.D. HOOKER.

[January 11th, 1844.]

Besides a general interest about the southern lands, I have been now ever since my return engaged in a very presumptuous work, and I know no one individual who would not say a very foolish one. I was so struck with the distribution of the Galapagos organisms, etc. etc., and with the character of the American fossil mammifers, etc. etc., that I determined to collect blindly every sort of fact, which could bear any way on what are species.

I have read heaps of agricultural and horticultural books, and have never ceased collecting facts. At last gleams of light have come, and I am almost convinced (quite contrary to the opinion I started with) that species are not (it is like confessing a murder) immutable. Heaven forfend me from Lamarck nonsense of a "tendency to progression," "adaptations from the slow willing of animals," etc.! But the conclusions I am led to are not widely different from his; though the means of change are wholly so. Ithink I have found out (here's presumption!) the simple way by which species become exquisitely adapted to various ends. You will now groan, and think to yourself, "on what a man have I been wasting my time and writing to." I should, five years ago, have thought so...

[The following letter written on February 23, 1844, shows that the acquaintanceship with Sir J.D. Hooker was then fast ripening into friendship. The letter is chiefly of interest as showing the sort of problems then occupying my father's mind:]

Dear Hooker, I hope you will excuse the freedom of my address, but I feel that as co-circum-wanderers and as fellow labourers (though myself a very weak one) we may throw aside some of the old-world formality...I have just finished a little volume on the volcanic islands which we visited. I do not know how far you care for dry simple geology, but I hope you will let me send you a copy. I suppose I can send it from London by common coach conveyance.

...I am going to ask you some MORE questions, though I daresay, without asking them, I shall see answers in your work, when published, which will be quite time enough for my purposes. First for the Galapagos, you will see in my Journal, that the Birds, though peculiar species, have a most obvious S. American aspect: I have just ascertained the same thing holds good with the sea-shells. It is so with those plants which are peculiar to this archipelago; you state that their numerical proportions are continental (is not this a very curious fact?) but are they related in forms to S. America. Do you know of any other case of an archipelago, with the separate islands possessing distinct representative species? I have always intended (but have not yet done so) to examine Webb and Berthelot on the Canary Islands for this object. Talking with Mr. Bentham, he told me that the separate islands of the Sandwich Archipelago possessed distinct representative species of the same genera of Labiatae: would not this be worth your enquiry? How is it with the Azores; to be sure the heavy western gales would tend to diffuse the same species over that group.

I hope you will (I dare say my hope is quite superfluous) attend to this general kind of affinity in isolated islands, though I suppose it is more difficult to perceive this sort of relation in plants, than in birds or quadrupeds, the groups of which are, I fancy, rather more confined. Can St. Helena be classed, though remotely, either with Africa or S. America?

>From some facts, which I have collected, I have been led to conclude that the fauna of mountains are EITHER remarkably similar (sometimes in the presence of the same species and at other times of same genera), OR that they are remarkably dissimilar; and it has occurred to me that possibly part of this peculiarity of the St. Helena and Galapagos floras may be attributed to a great part of these two Floras being mountain Floras. Ifear my notes will hardly serve to distinguish much of the habitats of the Galapagos plants, but they may in some cases; most, if not all, of the green, leafy plants come from the summits of the islands, and the thin brown leafless plants come from the lower arid parts: would you be so kind as to bear this remark in mind, when examining my collection.